ANSWER: POPULATION ECOLOGY
EXPLANATION:
The scientific field that deals with populations (i.e all individuals of a single species inhabiting a community) and how they interact with their extrinsic environment, is referred to as POPULATION ECOLOGY.
Thus, it's categorized under population ecology, because the research depicts how the number of individuals (beetles) in a population changes in time and the gross impact on their number decrease and loss of all the individuals (extinction) over a period of time.