By the time the United States purchased Alaska in 1867, decades of oppression and exposure to European diseases had taken their toll: The native population had dropped to just 2,500; the descendants of

By the time the United States purchased Alaska in 1867, decades of oppression and exposure to European diseases had taken their toll: The native population had dropped to just 2,500; the descendants of these survivors still make their home in the area today."

What does the phrase "taken their toll" mean here?

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