Answer:
nutrient, energy
Step-by-step explanation:
Organisms in an ecosystem are linked to their physical environment through the cycling of important nutrients and the transfer of energy from one trophic level to the other.
Important nutrients in the ecosystem are always in the state of flux, moving between living and non-living component of the environment with some processes that adds and removes these nutrients from the cycle.
Energy are also transferred from one trophic level to another within the ecosystem with a large amount being lost as heat.