Answer:
Option E is correct.
Step-by-step explanation:
The statement "Studying the fruit fly, a household nuisance but a time-honored experimental subject, has enabled the secrets of how embryos develop to begin to be unraveled by scientists" can be correctly written as
"The study of the fruit fly, a household nuisance but a time-honored experimental subject, has enabled scientist to begin to unravel the secrets of how embryos develop. "