Final answer:
The Commander of JTF-GNO is in charge of operating and defending Global Information Grid networks against cyber threats, as well as ensuring cybersecurity and information assurance for DoD network operations.
Step-by-step explanation:
The Commander (CO) of Joint Task Force-Global Network Operations (JTF-GNO) has two primary roles. Firstly, the CO is responsible for directing the operation and defense of Global Information Grid (GIG) networks against cyber threats. Secondly, the CO plays a pivotal role in ensuring the cybersecurity and information assurance of the Department of Defense's (DoD) network operations. These roles involve overseeing the security protocols, network operations, and other measures to protect against and respond to cyber incidents.