Final answer:
The correct word for the sentence is 'Although,' as it introduces a contrast between the length of the birds' journey and the small percentage of time they spend airborne.
Step-by-step explanation:
The correct word to complete the sentence is Although. The sentence should read: Although their journey takes four months, the birds spend only two to six per cent of the time actually in the air. The word 'Although' introduces a contrast between the duration of the birds' journey and the proportion of that time spent flying. It shows that in spite of taking a long time to complete their migration, birds are not flying continuously.