Final answer:
The exact number of tuskless elephants killed is not given in the provided references or question, hence an answer to this question cannot be provided. However, the materials focus on elephant conservation and the historical ivory trade.
Step-by-step explanation:
The question pertaining to the total number of elephants killed with meat intact but ivory naturally absent (tusklessness) for all areas surveyed is not directly answerable with the provided information.
There is no specific data included about the numbers of tuskless elephants killed that would allow for an accurate answer.
However, the provided reference materials discuss various aspects of elephant conservation, the history of ivory trade, and the impacts of hunting on wildlife populations.
To tackle the broader context, it's vital that strategic conservation efforts are made, such as establishing national parks and involving local communities in eco-tourism for sustainable development that can benefit both elephant populations and local people.