Final answer:
Other than DNA methylation, gene expression can be controlled at the DNA level through histone acetylation, transcription factors binding, and mRNA splicing.
Step-by-step explanation:
Gene expression can be controlled at the DNA level through several mechanisms. One such mechanism is histone acetylation, where the addition or removal of acetyl groups to histone proteins affects the accessibility of DNA for transcription. Another mechanism is the binding of transcription factors, which are proteins that bind to specific DNA sequences and either enhance or inhibit transcription. Additionally, mRNA splicing can also control gene expression by removing non-coding regions called introns from the pre-mRNA, allowing only the coding regions called exons to be translated into proteins.