Final answer:
The investigation showed that some of the QTLs had pronounced effects on the flowers' phenotypes.
Step-by-step explanation:
The results of Bradshaw and colleagues' investigation on the quantitative trait loci (QTL) of Mimulus species showed that some of the QTLs had pronounced effects on the flowers' phenotypes, indicating that they contribute significantly to the variation in flower traits. However, the differences in genotype for the majority of these QTLs explained small amounts of the variations in flower phenotypes for both species. Therefore, the correct answer is option 2) some of the QTLs did have pronounced effects on the flowers' phenotypes.