Final answer:
The perceived advantage linked to a disadvantageous trait, according to the handicap principle, is genetically linked to the sexual trait.
Step-by-step explanation:
According to the handicap principle, in an honest signal the perceived advantage linked to the disadvantageous trait is genetically linked to the sexual trait(b). This principle is part of the theory of sexual selection and suggests that only the fittest individuals can afford to have costly traits, such as a peacock's elaborate tail. While such traits may seem disadvantageous because they can increase visibility to predators or reduce mobility, they serve as honest signals of the individual's quality. Thus, females may prefer these males, who can pass strong genetic superiority to their offspring despite the apparent handicaps. This idea contrasts with direct benefits to the female or the trait not being inherited by offspring.