Final answer:
Common ancestors of cnidarians, flatworms, mollusks, and echinoderms evolved the trait of bilateral symmetry, despite echinoderms later exhibiting radial symmetry as adults.
Step-by-step explanation:
The trait that evolved in a common ancestor of cnidarians, flatworms, mollusks, and echinoderms is bilateral symmetry. Bilateral symmetry involves division through a midsagittal plane, creating two mirror images, which is a characteristic seen in the Bilateria clade. Even though adult echinoderms exhibit radial symmetry, their larval forms exhibit bilateral symmetry, indicating their evolutionary roots in bilaterally symmetrical ancestors.