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H+s are picked up from one side of the membrane, the __________, and are released to the other, into the __________ __________.

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Final answer:

H+ ions are picked up from one side of the membrane and released to the other side, either into the cytoplasm or extracellular fluid.

Step-by-step explanation:

The substances that are picked up from one side of the membrane are H+ ions.

These H+ ions are released to the other side of the membrane, into the cytoplasm or extracellular fluid.

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