Final answer:
Helmholtz successfully applied the principle of conservation of energy to living organisms.
Step-by-step explanation:
Helmholtz successfully applied the principle of conservation of energy to living organisms. This principle states that energy cannot be created or destroyed, but it can be transformed from one form to another. Helmholtz studied the energy transformations in the human body and developed the concept of energy conservation, which has been widely applied in the field of biology.
Learn more about Principle of conservation of energy in living organisms