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Before scientists learned how to make artificial fertilizers, what key process limited the amount of nitrates available to plants in most ecosystems?

Before scientists learned how to make artificial fertilizers, what key process limited the amount of nitrates available to plants in most ecosystems?
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Before scientists learned how to make artificial fertilizers, what key process limited the amount of nitrates available to plants in most ecosystems?

User Danielhrisca
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I believe rottening would be the answer. Rotting, dead plants and animal feaces make great fertilizers. 
User Marcello Miorelli
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