Final answer:
Research into the high mortality of marine iguanas due to ENSO could focus on the effects of climate change on their marine ecosystem and food sources, as well as increased predation from non-native species introduced to the Galápagos Islands.
Step-by-step explanation:
To understand why marine iguanas suffered a high mortality rate due to the abiotic effects of ENSO (El Niño Southern Oscillation), there are two possible research directions that could be pursued. Firstly, it is crucial to explore the impact of climate change on marine ecosystems, particularly how rapid temperature increases can cause species to become extinct before they have a chance to adapt. This includes examining how the iguanas' primary food sources are affected by increased sea temperatures and whether this leads to nutritional deficits.
Secondly, research should be conducted on the predatory pressures from non-native species, such as dogs and cats, which have been introduced to the Galápagos Islands and pose a threat to the iguanas. Understanding the increase in predation risk, combined with the stresses from ENSO, could explain the heightened mortality rates. This research would benefit from studies on the behavior and population dynamics of both the iguanas and their introduced predators during and after ENSO events.