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An advantage allowing the seed plant to greatly surpass the seedless vascular plants in colonizing the land was the evolution of:

User Pravchuk
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An advantage allowing the seed plant to greatly surpass the seedless vascular plants in colonizing the land was the evolution of air-dispersed sperm in pollen grains.Pollen is produced by the gymnosperms (and flowering plants as well) as a package for the dispersal of sperm.
User Kyler Love
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