How did Chargaff's results show that Levene's model was wrong? (See picture for table + diagram) Thank you!

In human DNA, for example, the four bases are present in these percentages: A=29.3% and C=20.0%; G=20.7% and T=28.4%. This strongly hinted towards the base pair makeup of the DNA.Chargaff is credited with disproving the Levene's widely accepted hypothesis that four bases in a dna are joined to each other..

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