The inversion of X-chromosome resulting in the movement of w+ gene to certain cell chromosomes.
Step-by-step explanation:
The mutation was first observed by Hermann Muller, in the year 1930.The eye colour variegation of flies is the result of unusual gene juxtaposition in certain cells through transposition. The translocation of gene in Drosophila the chromosome X gets inverted and places the w+ gene beside the heterochromatin.
Later, through the mutation the eye color becomes variegated and the gene is moved to certain cells of eyes, unlike the phenotype of red eye.