Answer:
Athelose and glucose.
Step-by-step explanation:
The ath operon most likely controlled due to the presence of athelose in small amount because ath operon are expressed only when the concentration of athelose in the bacterium is high. The smaller amount of athelose can be gained if glucose is present to the bacterium in sufficient amount or quantity so we can say that glucose quantity control the amount of athelose and it will control the ath operon.